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MicroRNA399 is induced to be expressed under phosphorus deficiency conditions and positively regulated by PHR1, which promotes phosphorus uptake and transport by inhibiting the mRNA expression of its target gene, the ubiquitin-conjugating E2 enzyme PHO2 , and then increasing protein expression of PHO1 and PHT1 in the downstream of PHO2 (Xiao et al., 2022b)

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